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When the Custom Operator Number is defined, if a caller presses '0' during the VoiceMail greeting playback, the call will be transferred to it. It is your prerogative as to whether you:
The caller can leave you a messsage first before pressing '0'. However, once transferred, the caller will no longer be able to leave a VoiceMail on TBB. Unless you have a separate VoiceMail system at your defined number, the caller will have to call you back.
Setting the Custom Operator Number
It should be specified preferably as 10 digits; long distance charges may apply to your TBB account if the number is prefixed with a "1".
1See Email Forwarding & Notifications (Phone, Pager, Cell Text Messaging) for an explanation of the other selections visible.